@article{fdi:010097484,
  title = {{N}ew fossil material of {C}helus ({T}estudines : {C}helidae) from the {M}iddle {M}iocene of {P}eru},
  author = {{C}adena, {E}. {A}. and {S}alas-{G}ismondi, {R}. and {Z}amora-{V}ega, {C}. and {F}lynn, {J}. and {A}ntoine, {P}. {O}. and {B}aby, {P}atrice and {P}{\'e}rez-{G}arcía, {A}.},
  editor = {},
  language = {{ENG}},
  abstract = {{W}e describe two fossil turtle shells of {C}helus colombiana from {M}iddle {M}iocene vertebrate localities in the {F}itzcarrald {A}rch and {I}quitos regions of {P}eru. {O}ne specimen represents one of the most complete fossil shells of the genus, also one of the largest known, measuring 84.6 cm in carapace length and estimated to have weighed approximately 65 kg. {M}orphological comparisons confirm synchronous consistency with other {C}. colombiana specimens from {C}olombia and {B}razil, with only minor variation in the gular and extragular scutes. {T}hese new records extend the known geographic distribution of {C}. colombiana to the southwestern margins of the {P}ebas wetland system and support a broader pattern of body size gigantism among {M}iocene {S}outh {A}merican turtles, likely driven by ecological connectivity rather than climatic factors. {H}owever, the absence of associated cranial material remains a major obstacle to fully resolving the taxon (c) 2026 {T}he {A}uthor(s). {P}ublished by {E}lsevier {B}.{V}. on behalf of {N}anjing {I}nstitute of {G}eology and {P}alaeontology.},
  keywords = {{S}outh {A}merica ; {N}eogene ; {P}ebas {F}ormation ; {T}estudines ; {P}leurodira ; paleobiogeography ; {PEROU}},
  booktitle = {},
  journal = {{P}alaeoworld},
  volume = {35},
  numero = {5},
  pages = {201135 [10 p.]},
  ISSN = {1871-174{X}},
  year = {2026},
  DOI = {10.1016/j.palwor.2026.201135},
  URL = {https://www.documentation.ird.fr/hor/fdi:010097484},
}