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    <titleInfo>
      <title>Evidence for non-random co-occurrences in a white shark aggregation</title>
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    <abstract>Groups or aggregations of animals can result from individuals being attracted to a common resource or because of synchronised patterns of daily or seasonal activity. Although mostly solitary throughout its distribution, white sharks (Carcharodon carcharias) seasonally aggregate at a number of sites worldwide to feed on calorie-rich pinnipeds. At the Neptune Islands, South Australia, large numbers of white sharks can be sighted throughout the year, including during periods of low seal abundance. We use a combination of photo-identification and network analysis based on co-occurrence of individuals visiting the site on the same day to elucidate the population structure and aggregatory behaviour of Australia's largest aggregation of sub-adult and adult white sharks. We photo-identified 282 sharks (183 males, 97 females, 2 unknown) over a 4.5-year period (June 2010-November 2014) and found that white sharks did not randomly co-occur with their conspecifics, but formed four distinct communities. Tendency to co-occur varied across months with males co-occurring with more individuals than females. Sex-dependent patterns of visitation at the Neptune Islands and resulting intraspecific competition likely drive the observed community structure and temporal variability in co-occurrences. This study provides new insights into the aggregatory behaviour of white sharks at a seal colony and shows for the first time that white shark co-occurrence can be non-random.</abstract>
    <targetAudience authority="marctarget">specialized</targetAudience>
    <subject>
      <topic>Social network analysis</topic>
      <topic>Gregariousness</topic>
      <topic>Social behaviour</topic>
      <topic>Aggregation</topic>
      <topic>Photo-ID</topic>
      <topic>Carcharodon carcharias</topic>
    </subject>
    <subject authority="local">
      <geographic>AUSTRALIE</geographic>
      <geographic>NEPTUNE ILES</geographic>
    </subject>
    <classification authority="local">034</classification>
    <classification authority="local">020</classification>
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      <titleInfo>
        <title>Behavioral Ecology and Sociobiology</title>
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      <part>
        <detail type="volume">
          <number>73</number>
        </detail>
        <detail type="volume">
          <number>10</number>
        </detail>
        <extent unit="pages">
          <list> art. 138 [12 p.]</list>
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      <originInfo>
        <dateIssued>2019</dateIssued>
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      <identifier type="issn">0340-5443</identifier>
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    <identifier type="uri">https://www.documentation.ird.fr/hor/fdi:010077094</identifier>
    <identifier type="doi">10.1007/s00265-019-2745-1</identifier>
    <identifier type="issn">0340-5443</identifier>
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