@article{fdi:010073749,
  title = {{I}ntegration of nuclear and mitochondrial gene sequences and morphology reveals unexpected diversity in the forest cobra ({N}aja melanoleuca) species complex in {C}entral and {W}est {A}frica ({S}erpentes : {E}lapidae)},
  author = {{W}uster, {W}. and {C}hirio, {L}. and {T}rape, {J}ean-{F}ran{\c{c}}ois and {I}neich, {I}. and {J}ackson, {K}. and {G}reenbaum, {E}. and {B}arron, {C}. and {K}usamba, {C}. and {N}agy, {Z}. {T}. and {S}torey, {R}. and {H}all, {C}. and {W}uster, {C}. {E}. and {B}arlow, {A}. and {B}roadley, {D}. {G}.},
  editor = {},
  language = {{ENG}},
  abstract = {{C}obras are among the most widely known venomous snakes, and yet their taxonomy remains incompletely understood, particularly in {A}frica. {H}ere, we use a combination of mitochondrial and nuclear gene sequences and morphological data to diagnose species limits within the {A}frican forest cobra, {N}aja ({B}oulengerina) melanoleuca. {M}itochondrial {DNA} sequences reveal deep divergences within this taxon. {C}ongruent patterns of variation in mt{DNA}, nuclear genes and morphology support the recognition of five separate species, confirming the species status of {N}. subfulva and {N}. peroescobari, and revealing two previously unnamed {W}est {A}frican species, which are described as new: {N}aja ({B}oulengerina) guineensis sp. nov. {B}roadley, {T}rape, {C}hirio, {I}neich & {W}uster, from the {U}pper {G}uinea forest of {W}est {A}frica, and {N}aja ({B}oulengerina) savannula sp. nov. {B}roadley, {T}rape, {C}hirio & {W}uster, a banded form from the savanna-forest mosaic of the {G}uinea and {S}udanian savannas of {W}est {A}frica. {T}he discovery of cryptic diversity in this iconic group highlights our limited understanding of tropical {A}frican biodiversity, hindering our ability to conserve it effectively.},
  keywords = {{I}ntegrative taxonomy ; {A}frica ; {N}aja melanoleuca ; {N}aja guineensis sp nov. ; {N}aja savannula sp nov. ; {E}lapidae ; systematics ; {AFRIQUE} {CENTRALE} ; {AFRIQUE} {DE} {L}'{OUEST} ; {ZONE} {TROPICALE}},
  booktitle = {},
  journal = {{Z}ootaxa},
  volume = {4455},
  numero = {1},
  pages = {68--98},
  ISSN = {1175-5326},
  year = {2018},
  DOI = {10.11646/zootaxa.4455.1.3},
  URL = {https://www.documentation.ird.fr/hor/fdi:010073749},
}